The famous grizzly bear mother, a longtime icon of the Greater Yellowstone Ecosystem, has died after being struck by a vehicle in Wyoming.

Grizzly 399 had a yearling cub with her when she was struck by the vehicle. The cub's whereabouts are unknown, but there is no evidence it was involved.

She lived to 28, roaming the Yellowstone area with her many offspring while becoming a favorite among both residents and visitors.